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It was called Ephemerella hecuba subgenus Timpanoga until about 1990. The name Ephemerella was dropped as a genus and Timpanoga was applied. The following paper (1959) gives some additional information on the nymph. http://www.ephemeroptera-galactica.com/pubs/pub_a/puballenr1959p51.pdf

Steve, I can't remember. It's been years since I observed them and I couldn't find any in a cursory search of specimens. I would assume they have at least a limited amount of movement. I did find a photo of a nymph covered in debris though. https://joycegross.com/images.php?img=2222_0939

I imagine that the "spikeyness" would help to deter predators, along with it's width. In the creek where I found them the primary predators would be Caifornia roach, the fry of steelhead and coho, occasionally a steelhead that stayed over a year, sculpins, and dragonfly larvae. I suppose that...

They're the operculae {singular-operculum} and gills. The operculum is on top of the gills and made of a horny substance, The gills are underneath, protected from silt and other detritus. The operculae are probably an adaptation due to the nymph almost always sitting on the bottom in slackwater...

Hey Roger, Thanks for the compliment. You've certainly looked at many photos of macroinvertebrates. However I believe that almost anybody can take good photos of them with the cameras of today.

The ones on top are operculae. They protect the gills from the silt and other irritants. The gills are underneath these. You can see a little bit of the gills poking out in the bottom picture.

Usually found in pools or slackwater areas. The nymphs often have a coating of diatoms and silt on them. Size ranges from about 7-10 mm {not including tails) at maturity. These were collected in Mill Creek (tributary to the Russian River).

These nymphs are common in Palmer Creek and Felta Creek, tributaries to Mill Creek which is a tributary to the Russian River. Usually found in glides and fast water.

Looks like Skwala. https://theeveninghatch.com/the-invisible-imaginary-skwala https://bugguide.net/node/view/258802/bgimage

Actually, they were modeled after wasp larvae. Here's one in action.

These Ecdyonurus nymphs and their parasites were found in Mill Creek, a tributary of the Russian River. The Symbiocladius parasites are members of Diptera (fly family). I've only found them on Ecdyonurus nymphs although they are also found on Rhithrogena and Paraleptophlebia nymphs as well. They...

Not really, they're found in the same area, they all have the same sharp projections on the femur of the forelegs and the same sharp projections on the dorsal side of the abdomen. A lot of ephemerellids have color patterns that vary widely among individuals.

Drunella flavilinea nymphs are common in the Russian River from mid-February through May. They prefer areas of fast water and cobbled bottoms. There is quite a bit of color variation in these nymphs.

Crane fly, I think. https://news.arizona.edu/story/whats-all-crane-flies

I decided to order the book again after reading Steelhead Dave's account. There were some other articles and a youtube video that were interesting too. https://hakaimagazine.com/features/the-halibut-hook-revival/ http://arcadianabe.blogspot.com/2013/03/making-bentwood-halibut-hooks.html

Ensatina eschscholtzii https://amphibiaweb.org/cgi/amphib_query?where-genus=Ensatina&where-species=eschscholtzii&account=amphibiaweb Female Male Juveniles Female in defensive pose Female releasing toxin. Milky white on head and tail. Closeup of toxin Male releasing toxin Male...

Do you know what kind of salamander it is?

I remember having this book back in the late 70s or early 80s. It was (is) a great book. I think I'll try ordering it again.